Exploring the Binding and ATPase Activity of Msh2-Msh6 with Four Way Junction DNA

Amy Du
Amy Du

Amy is a rising senior originally from Castro Valley, CA. She attended high school at The College Preparatory School in Oakland. She is an MB&B and Economics major. After college, she plans to take a few years off before applying to medical school. Outside of school, she enjoys playing volleyball, cooking and baking (and eating), and watching criminal psychology videos on Youtube.

Abstract: Msh2-Msh6 is a eukaryotic ATP-dependent mismatch repair (MMR) protein that plays a key role in maintaining the fidelity of the genetic code. The protein recognizes and initiates repair for single base pair mismatches and short insertion/deletion loops (IDL). Fluorescence in solution assays have revealed that Msh2-Msh6 binds with the highest affinity to G:T mismatch DNA. Consequently, deficiencies in Msh2-Msh6 function are implicated in cancer etiology and specifically that of hereditary nonpolyposis colorectal cancer. Interestingly, fluorescence in-solution assays have revealed that Msh2-Msh6 binds to Holliday Junction DNA with 1:1 stoichiometry and a binding affinity of 12.9±1.3nM, similar to its affinity for G:T mismatch. However, the functional importance and details of this interaction are uncharacterized. We employ ATPase activity assays and equilibrium binding assays to understand the function of Msh2-Msh6’s interaction with Holliday Junction DNA; namely, we conduct experiments with G:T heteroduplex DNA and J3 Holliday Junction to establish control measurements. We follow with experiments conducted with J3 junction DNA that includes a G:T mismatch incorporated at various positions to probe the ATP hydrolysis and binding activity of Msh2-Msh6 in the presence of two favorable binding targets.

Video: https://wesleyan.zoom.us/rec/share/JloEfUJ2LNODVl0i1R2y6Xoxt3cyMd-fTCqH1nTE5oJcHbWbmoYUBxzclRrZ4W0X.D9RNV3s80yf07Mtf 

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